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The Pennsylvania State University is a public state-related land-grant research university with campuses and facilities throughout Pennsylvania.
Whitehead Institute for Biomedical Research is a leading, nonprofit research and educational institution that has defined the cutting edge of biomedical science, creating a legacy of research excellence and academic eminence since 1982. Wholly independent in its governance, finances and research programs, Whitehead shares a teaching affiliation with Massachusetts Institute of Technology, offering the intellectual, collegial and scientific benefits of a leading research university.
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